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Solution Carnivorous Plants Insects To Obtain Nitrogen Jawaban Trial Environment TypesMost plants take up and water and nutrients from the CO2 from the air through the shoot system; soil through the root system Concept 37.1: Soil contains a Plants obtain most of their water and minerals from the Living organisms play an important role in these soil layers The basic physical properties of soil are living, complex ecosystem; upper layers of soil; -Texture - Composition Soil Texture Soil particles are classified by size: from largest to smallest, they are called Soil is stratified into layers called - Topsoil consists of which is decaying organic material After a heavy rainfall, water drains from the larger spaces in the soil, but smaller spaces retain water because of sand, silt, and clay; soil horizons; mineral particles, living organisms, and humus, its attraction to clay and other particles Loams are the most and contain equal amounts of Sandy soils dont retain enough; clay soils retain The most fertile top-soils typically have pores containing fertile top soils; sand, silt, and clay water to support plant growth; too much about half water and half air Topsoil Composition A soils composition refers to its inorganic (mineral) and organic chemical components Inorganic Components Cations (for example K, Ca2, and Mg2) adhere to; this prevents them from through percolating groundwater During cation exchange, cations are displaced from, particularly H -- negatively charged soil particles; leaching out of the soil soil particles by other cations Displaced cations enter the soil solution and can be taken Negatively charged ions with soil particles and can be up by plant roots do not bind; lost from the soil by leaching Organic Components Humus builds a crumbly soil that It also increases the soils capacity to and - retains water but is still porous; exchange cations; serves as a reservoir of mineral nutrients Topsoil contains - These organisms help to and bacteria, fungi, algae, other protists, insects, earthworms, nematodes, and plant roots; decompose organic materia; and mix soil Soil Conservation and Sustainable Agriculture Fertilization, the addition of, combats the reduction in yields experienced when crops are cultivated in the same location over many seasons Soil management, by fertilization and other practices, In contrast with natural ecosystems, - mineral nutrients to the soil; allowed for sedentary agriculture and the formation of villages; agriculture depletes the mineral content of soil, taxes water reserves, and encourages erosion The American Dust Bowl of the 1930s resulted from At present, of the worlds farmland has reduced productivity due to soil mismanagement The goal of sustainable agriculture is to use farming methods that are soil mismanagement; 30 conservation-minded, environmentally safe, and profitable Irrigation Irrigation is a huge drain on when used for farming in arid regions For example, of global freshwater use is devoted to agriculture The primary source of irrigation water is underground water reserves The depleting of aquifers can result in Irrigation can lead to salinization, the concentration of water resources; 75 called aquifers; land subsidence, the settling or sinking of land salts in soil as water evaporates Salinization reduces the diminishes Drip irrigation by slowly water potential of the soil; water uptake by plants reduces water use and salinization; releasing water directly at the root zone Fertilization Soils can become depleted of nutrients as plants and the nutrients they contain are harvested Fertilization replaces mineral nutrients that have been lost from the soil Commercial fertilizers are enriched in Excess minerals are often leached from the Organic fertilizers are composed of manure, fishmeal, or compost They gradually release nitrogen (N), phosphorus (P), and potassium (K) soil and can cause algal blooms in lakes N, P, and K as they decompose Adjusting Soil pH Soil pH affectsand the Cations are more available in, as H ions displace mineral cations from clay particles --The availability of For example, at pH 8, plants can absorb calcium but not iron At soil pH 5 or lower, toxic aluminum ions (Al3) become more soluble Uptake of Al3 cation exchange; chemical form of minerals slightly acidic soil; different minerals varies with Ph stunts root growth and prevents calcium uptake Controlling Erosion: Water and wind erosion are major causes of - Erosion can be reduced by - No-till agriculture also reduces erosion by creating topsoil removal and loss of soil nutrients planting trees as windbreaks, terracing hillside crops, and cultivating in a contour pattern furrows for seeds and fertilizer with minimal soil disturbance Phytoremediation Some areas are unfit for agriculture due to or Phytoremediation is a biological, nondestructive technology that Plants capable of extracting soil pollutants are contamination of soil; groundwater with toxic pollutants reclaims contaminated areas grown and are then disposed of safely Concept 37.2: Plant roots absorb essential elements from the soil Water, air, and soil minerals all contribute to plant growth - of a plants fresh mass is water - 96 of a plants dry mass is from - 4 of a plants dry mass is 80-90; CO2 assimilated into carbohydrates during photosynthesis inorganic substances from soil Essential Elements Not all of the more than 50 chemical elements identified There are, chemical elements required for a plant to Researchers use to determine which chemical elements are essential in plants are essential; 17 essential elements complete its life cycle and reproduce hydroponic culture Nine of the essential elements are called because plants require them in The macronutrients are contributes the most to plant growth and crop yields macronutrients; relatively large amounts carbon, oxygen, hydrogen, nitrogen, phosphorus, sulfur, potassium, calcium, and magnesium Nitrogen The remaining eight are called because plants need them in very small amounts The micronutrients are Plants with also require sodium Micronutrients function as - micronutrients; chlorine, iron, manganese, boron, zinc, copper, nickel, and molybdenum C4 and CAM photosynthetic pathways cofactors, nonprotein helpers in enzymatic reactions Symptoms of Mineral Deficiency Symptoms of mineral deficiency depend on the Deficiency of a mobile nutrient usually affects Deficiency of a less mobile nutrient usually The most common deficiencies are those of nutrients function and mobility within the plant older organs more than young ones affects younger organs more than older ones nitrogen, potassium, and phosphorus Improving Plant Nutrition by Genetic Modification Plants can be to improve genetically engineered; nutrition and fertilizer usage Resistance to Aluminum Toxicity: Aluminum in acidic soils and -The introduction of bacterial genes into plant genomes can cause damages roots; greatly reduces crop yields plants to secrete acids that bind to and tie up aluminum Smart Plants -- Smart plants inform the grower of a before damage has occurred - A blue tinge indicates when these plants need nutrient deficiency; phosphorus-containing fertilizer Concept 37.3: Plant nutrition often involves Plants have a mutualistic relationships with Dead plants provide energy needed by relationships with other organisms soil bacteria and fungi soil-dwelling microorganisms Secretions from support a wide variety of Many mutually beneficial relationships occur between species from living roots; microbes in the near-root environment; different kingdoms or domains Fungus-Bacterium A lichen is a mutualistic association between a Animal-Bacterium: Most puffer fish form a mutualistic association with a Plant-Bacterium - The floating fern Azolla forms mutualistic associations with fungus and a photosynthetic partner; bacterium that produces a nerve toxin called tetrodotoxin nitrogen-fixing cyanobacteria Animal-Fungus - Leaf-cutter ants harvest leaves to provide nutrition to Plant-Fungus - Most plant species are associated Plant-Animal - Some species of Acacia plants provide fungal gardens in their nests with mycorrhizal fungi nourishment to ants that protect them from predators and competitors Bacteria and Plant Nutrition - A variety of soil bacteria play roles in plant nutrition Rhizobacteria - The is the rhizosphere Rhizobacteria are or in the rhizosphere Some rhizobacteria are; others are endophytes, layer of soil closely surrounding the plants roots bacteria that live in close association with plant roots free living; nonpathogenic bacteria that live between the cells of host plant tissues Endophytes and rhizobacteria depend on and, in return, help to enhance plant growth by - nutrients secreted by plant cells; Producing antibiotics that protect roots from disease Absorbing toxic metals or increasing nutrient availability Converting nitrogen into forms usable by the plant Producing chemicals that stimulate plant growth Bacteria in the Nitrogen Cycle Nitrogen can be an important limiting nutrient for plant growth Plants can absorb nitrogen as either Some soil nitrogen derives from, but most comes from the NO3- or NH4 inorganic sources; activity of soil bacteria Plants mainly acquire nitrogen as Soil NO3- is formed in a two-step process called Nitrifying bacteria to Different nitrifying bacteria oxidize nitrate (NO3-); nitrification oxidize ammonia (NH3); nitrite (NO2-) NO2- to NO3- Plant enzymes convert, which is incorporated into organic compounds -- Some soil nitrogen is lost to the atmosphere when denitrifying bacteria Plants can also use NH4, which is acquired NO3- to ammonium (NH4) convert NO3- to gaseous nitrogen (N2) through two separate processes Nitrogen-fixing bacteria convert; NH3 picks up an Ammonifying bacteria break down N2 into NH3 H in the soil solution and forms NH4 dead organic compounds and release NH4 Nitrogen-Fixing Bacteria: A Closer Look (between Rhizobium and Legumes) Nitrogen is abundant in the atmosphere but unavailable to plants because of the -- Nitrogen fixation is the conversion of nitrogen Some nitrogen-fixing bacteria are triple bond between atoms in N2 from N2 to NH3 free-living in the soil; others are rhizospheric nitrogen-fixing Rhizobium (root-living) bacteria provide some plant species (e.g., legumes) with a - Along a legumes roots are swellings called infected by Rhizobium bacteria Inside the root nodule, Rhizobium bacteria assume a form called formed by the root cell- source of fixed nitrogen; nodules composed of plant cells bacteroids, which are contained within vesicles The plant obtains fixed nitrogen from, and Rhizobium Each legume species is associated with a particular strain of The development of a nitrogen-fixing root nodule depends on chemical dialogue between Rhizobium; obtains sugar and an anaerobic environment Rhizobium bacteria Rhizobium bacteria and root cells of their specific plant hosts Crop rotation takes advantage of the agricultural benefits of symbiotic nitrogen fixation A nonlegume such as maize is planted - Instead of being harvested, the legume crop is often plowed under to decompose as - Nonlegumes such as alder trees and certain tropical grasses are associated with nitrogen-fixing bacteria - Rice paddies often contain an aquatic fern that has mutualistic cyanobacteria that fix nitrogen one year, and the next year a legume is planted to restore the concentration of fixed nitrogen in the soil; green manure Mycorrhizae are mutualistic The host plant provides the fungus with a The fungus increases the surface area for associations of fungi and roots steady supply of sugar water uptake by the host plant and supplies the plant with mineral nutrients from the soil Mycorrhizal fungi secrete They also secrete antibiotics to help protect the growth factors that stimulate root growth and branching; plant from soil pathogens Mycorrhizae and Plant Evolution Early land plants would have encountered harsh conditions 400 to 500 million years ago These plants lacked the ability to extract essential nutrients from the soil, The mycorrhizal association between early land plants and fungi allowed both while fungi were unable to produce carbohydrates; to exploit the terrestrial environment Types of Mycorrhizae - Mycorrhizal associations consist of two major types: Ectomycorrhizae -- Arbuscular mycorrhizae In ectomycorrhizae, the - These hyphae form a network in the - Ectomycorrhizae occur in about 10 of mycelium of the fungus forms a dense sheath, or mantle, over the surface of the root apoplast, do not penetrate the root cells plant families, most of which are woody; In arbuscular mycorrhizae, microscopic fungal hyphae These mycorrhizae penetrate the Hyphae form branched Arbuscular mycorrhizae occur in about 85 of plant species, including most crops About 5 of plant species do not form mycorrhizal associations extend into the root cortex; cell wall but not the plasma membrane arbuscules within cells; these are important sites of nutrient transfer Agricultural and Ecological Importance of Mycorrhizae Farmers and foresters often to promote formation of mycorrhizae - Some invasive exotic plants disrupt interactions - For example, garlic mustard slows growth of other plants by preventing the growth of mycorrhizal fungi inoculate seeds with fungal spores; between native plants and their mycorrhizal fungi Epiphytes, Parasitic Plants, and Carnivorous Plants Some plants have nutritional adaptations that use other organisms in Three unusual adaptations are - nonmutualistic ways Epiphytes - Parasitic plants - Carnivorous plants Epiphytes An epiphyte grows on Epiphytes do not tap into hosts for sustenance another plant and obtains water and minerals from rain Parasitic Plants Parasitic plants absorb Some species also Some species parasitize the sugars and minerals from their living host plant photosynthesize, but others rely entirely on the host plant for sustenance mycorrhizal hyphae of other plants Carnivorous Plants: Carnivorous plants have adaptations for - They are trapping insects and other small animals; 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